172
CHECKERED BEETLES AND ALLIES
CLEROIDEA
T
he Cleroidea is a medium-sized superfamily
within Polyphaga, comprising just over 10,000
known species worldwide, in 13 families. However,
as with many beetle superfamilies, the species are
unevenly spread across the families. Four families,
Phloiophilidae, Metaxinidae, Phycosecidae, and
Acanthocnemidae, each has only a single genus with
one to four species, while Cleridae and Melyridae,
covered separately below, each include several
thousand and 90 percent of the species between
them. The third largest family of the Cleroidea is the
Trogossitidae, known as “bark-gnawing beetles,”
with about 600 species in 50 genera. Many of these
are predators found in the tunnels of bark beetles
and other wood-boring insects. Some are useful
biological control agents of forestry pests. One
species—“the cadelle,” Tenebroides mauritanicus—
was spread all over the world with stored products,
though it probably often eats other stored-product
insects rather than the crops themselves.
CLEROIDEA—Checkered Beetles and Allies
superfamily
Cleroidea
known species
10,500
distribution
Worldwide except Antarctica
habitat
Most habitats, particularly forests. Some
species in stored products
size
2–50 mm
diet
Many species are predators, while others
are flower feeders or feed on fungi
notes
The only member of the Phloiophilidae, the
European Phloiophilus edwardsii, is a small
beetle 2–3 mm long, attractively marked in
brown and black. It occurs on the fungus
Peniophora quercina growing on the twigs